Ett kunskapscentrum för Sveriges arter och naturtyper
Ett kunskapscentrum för Sveriges arter och naturtyper

Per Alström's research

Per Alström in Arunachal Pradesh 2009. Photo: Shashank Dalvi

Research

Taxonomy of passerine birds

The number of recognized bird species in the world has increased by 20% in the last half century. Most of this increase is due to improved taxonomic knowledge. This stems mainly from application of new methods, especially analyses of DNA and vocalizations. Together with a number of co-workers, I study the taxonomy of various groups of passerine birds, mainly different “warblers”, using multiple lines of evidence, such as morphology, DNA, vocalizations, other behaviours, ecology, geographical distributions etc.

Background

New methods – DNA and vocalizations

Traditionally, taxonomy has been based on studies of morphology: forms with a certain (subjective) degree of similarity have been treated as subspecies of the same species. However, in recent years,studies of DNA have frequently revealed that forms that are treated as conspecific are highly divergent, and sometimes not even each other’s closest relatives. Moreover, research on vocalizations has shown that many forms treated as subspecies of the same species have highly distinct songs, and do not respond to songs of other forms that are considered to belong to the same species. Especially vocal studies have also frequently revealed the existence of previously unknown so-called “cryptic species” (i.e. different species that are very similar in appearance).

Exploding species

Studies involving DNA and vocalizations, often in combination with morphological (plumage, structure), behavioural, ecological and distributional data, have shown that certain forms warrant recognition as separate species rather than as subspecies of the same species.The increase in the number of recognized species due to recent research based on these modern taxonomic methods has often been dramatic. For example, in the family Phylloscopidae (comprising the genera Phylloscopus and Seicercus), which is one of the groups that I have been studying, the number of species has almost doubled in the last 25 years.

Research

Most of the taxonomic work that I am involved with concerns passerine birds, especially various groups of “warblers”, larks, pipits and wagtails. My co-workers and I use as many different lines of evidence as possible in evaluations of taxonomic status. The research has revealed many previously unrecognized species, including a few that were previously unknown to science. Some examples:

 Arctic Warblers -- three anciently separated species [Vocalizations]

 "Golden-spectacled Warblers" -- a complex of cryptic species [Vocalizations]

 Spotted Bush Warblers -- three species instead of one [Songs]

 Yellowish-bellied Bush Warblers -- two distinct species [Vocalizations]

Systematics of passerine birds

Systematics is the science of evolutionary relationships (phylogeny) among organisms. Until recently, relationships among birds were inferred based on similarities in external and anatomical features. However, in recent years, analyses of DNA have revolutionized these studies. Most of the traditionally deduced relationships have been confirmed by the new methods. However, DNA analyses have also rebutted many of the old opinions and revealed previously completely unexpected relationships. Together with a number of co-workers in several different countries, I study systematics of passerine birds using DNA sequences.

Background

Closely related species look similar – but not always

In general, closely related species are more similar in appearance to each other than to more distantly related species. However, this needs not be the case. Species that are not closely related might evolve similar features because their live under similar conditions (e.g., different groups of swimming birds have independently evolved webbed feet, and desert birds are generally paler, more “sand-coloured” than relatives in less dry areas). This is called parallel or convergent evolution. Moreover, in some groups of birds, some species have evolved strikingly different features from their close relatives, obscuring their relationships.

Secrets revealed by DNA

Analyses of DNA have revolutionized phylogenetic studies. DNA sequences have become easier and cheaper to obtain, and new statistical methods and more powerful computers have enabled more rigorous assessments of relationships. Although most of the traditionally deduced relationships have been confirmed by the new methods, molecular studies have revealed multiple examples of misconceived relationships due to highly complex morphological evolution.

Research

My colleagues and I use DNA sequence data, usually from both mitochondrial and nuclear DNA, to infer relationships of passerine birds. Many studies concern species within a genus, whereas other studies deal with relationships among genera within a family or among families in a superfamily. Most of these studies have discovered surprising relationships. Some examples:

 Unexpected relationships within the family Acrocephalidae

 Unexpected relationships and complex morphological evolution in the family Cettiidae

 Phylogeny of the family Locustellidae reveals multiple unexpected relationships

Various

Rediscovery of the enigmatic Blackthroat Luscinia obscura 

In June 2011, together with Dr Gang Song, Institute of Zoology, Chinese Academy of Sciences, Beijing, Dr Xue-bin Gao, Shaanxi Institute of Zoology, Xian, Mr Huisheng Gong, Foping National Nature Reserve, and Mr Yong-wen Zhang, Changqing National Nature Reserve, I rediscovered the breeding area for the poorly known Blackthroat Calliope obscura (Luscinia obscura), in the Qinling mountains, Shaanxi province, north central China. Seven singing males were observed in Foping and seven more in Changqing National Nature Reserves – which almost equals the total number of individuals observed of this species since its discovery in the late 19th century. Nearly all of the birds were on mountain slopes at 2400–2500 meters above sea level in large, dense expanses of bamboo in open coniferous and mixed coniferous-broadleaved forest.

 Adult male Blackthroat Luscinia obscura, Foping National Nature Reserve, Shaanxi, China, June 2011. © Per Alström

 Song of male Blackthroat Luscinia obscura, Foping National Nature Reserve, Shaanxi, China, 8 June 2011. © Per Alström 

 Song of male Blackthroat Luscinia obscura, Changqing National Nature Reserve, Shaanxi, China, 15 June 2011. © Per Alström

Publications

Papers in international peer-reviewed journals

The papers for which pdfs are available have 'pdf' written after the journal information. Some of these can be accessed by clicking on 'pdf'. To request a pdf that is not available here, please write to per.alstrom@slu.se. The open access papers can be accessed by clicking on 'Open access'.

Wang, W., Dai, C. Alström, P., Zhang, C., Qu, Y., Li, S.-h., Yang, X., Zhao, N.Song, G. & Lei, F. 2014 (in press). Past hybridization between two East Asian long-tailed tits (Aegithalos bonvaloti and A. fuliginosus). Frontiers in Zoology. [Abstract]

Lei, F., Qu, Y., Song, G., Alström, P. & Fjeldså, J. 2014 (in press). The potential drivers in forming avian biodiversity hotspots in the East Himalaya-Mountains of Southwest China. Integrative Zoology.

Price, T.D., Hooper, D.M., Buchanan, C.D., Johansson, U.S., Tietze, D.T., Alström, P., Olsson, U., Ghosh-Harihar, M., Ishtiaq, F., Gupta, S.K., Martens, J.E., Harr, B., Singh, P. & Mohan, D. 2014. Niche filling slows the diversification of Himalayan songbirds. Nature. (http://dx.doi.org/10.1038/nature13272) [Abstract] [English popular account] [Svensk populärvetenskaplig sammanfattning] pdf

Alström, P., Hooper, D.M., Liu, Y., Olsson, U., Mohan, D., Gelang, M., Hung, L.M., Zhao, J., Lei, F. & Price, T.D. 2014. Discovery of a relict lineage and monotypic family of passerine birds. Biology Letters vol. 10 no. 3 20131067. (http://dx.doi.org/10.1098/rsbl.2013.1067) [Abstract] [Figure 2pdf

Song, G., Alström, P., Zhang, Y., Gao, X., Gong, H., Holt, P.I., Qing, Q., Yin, Z. & Lei, F. 2013. Rediscovery of an enigmatic Chinese passerine, the Blackthroat Calliope obscura: plumage, vocalizations, distribution, habitat choice, nesting and conservation. Journal of Ornithology. (http://dx.doi.org/10.1007/s10336-013-1009-5) [Abstract] pdf

Alström, P., Song, G., Zhang, R., Gao, X., Holt, P.I., Olsson, U. & Lei, F. 2013. Taxonomic status of Blackthroat Calliope obscura and Firethroat C. pectardens. Forktail 29: 94–99. [Abstract] pdf

Alström, P., Barnes, K.N., Olsson, U., Barker, F.K., Bloomer, P., Khan, A.A., Qureshi, M.A., Guillaumet, A., Crochet, P.-A. & Ryan, P.G. 2013. Multilocus phylogeny of the avian family Alaudidae (larks) reveals complex morphological evolution, non-monophyletic genera and hidden species diversity. Molecular Phylogenetics and Evolution 69: 1043–1056 http://dx.doi.org/10.1016/j.ympev.2013.06.005 [Abstract] pdf

Xia, C., Liu, J., Alström, P., Wu, Q. & Zhang, Y. 2013. Is the soft song of the Brownish-flanked Bush Warbler an aggressive signal? Ethology 119: 653–661. http://dx.doi.org/10.1111/eth.12104 [Abstract] pdf

Alström, P., Olsson, U & Lei, F. 2013. A review of the recent advances in the systematics of the avian superfamily Sylvioidea. Chinese Birds 4: 99–131. http://dx.doi.org/10.5122/cbirds.2013.0016 pdf

Olsson, U. & Alström, P. 2013. Molecular evidence suggests that the enigmatic Sulawesi endemic Geomalia heinrichi belongs in the genus Zoothera (Turdidae, Aves). Chinese Birds 4: 155–160. http://dx.doi.org/10.5122/cbirds.2013.0015 pdf

Spottiswoode, C.N., Olsson, U., Mills, M.S.L., Cohen, C., Francis, J.E., Toye, N., Hoddinott, D., Dagne, A., Wood, C., Donald, P.F., Collar, N.J., Alström, P., 2013. Rediscovery of a long-lost lark reveals the conspecificity of endangered Heteromirafra populations in the Horn of Africa. Journal of Ornithology 154: 813–825. http://dx.doi.org/10.1007/s10336-013-0948-1 [Abstract] pdf

Olsson, U., Yosef, R & Alström, P. 2013. Assessment of species limits in African ‘brown buntings’ (Emberiza, Passeriformes, Aves) based on mitochondrial and nuclear sequence data. Ibis 155: 534–543. http://dx.doi.org/10.1111/ibi.12044 [Abstract] pdf

Xing, X.Y., Alström, P., Yang, X.J. & Lei, F.M. 2013. Recent northward range expansion promotes song evolution in a passerine bird, the Light-vented Bulbul. Journal of Evolutionary Biology 26: 867–877. doi: 10.1111/jeb.12101. [Abstract] pdf

Olsson, U., Leader, P.J., Carey, G.J., Khan, A.A., Svensson, L. & Alström, P. 2013. New insights into the intricate taxonomy and phylogeny of the Sylvia curruca complex. Molecular Phylogenetics and Evolution 6 7: 72–85. http://dx.doi.org/10.1016/j.ympev.2012.12.023. [Abstract] pdf

Olsson, U., Irestedt, M.,  Sangster, G., Ericson, P.G.P. & Alström, P. 2013. Systematic revision of the avian family Cisticolidae based on a multi-locus phylogeny of all genera. Molecular Phylogenetics and Evolution 66: 790–799. http://dx.doi.org/10.1016/j.ympev.2012.11.004. [Abstract] pdf

Fregin, S., Haase, M., Olsson, U. & Alström, P. 2012. New insights into family relationships within the avian superfamily Sylvioidea (Passeriformes) based on seven molecular markers. BMC Evolutionary Biology 12: 157. DOI: 10.1186/1471-2148-12-157 [Open access] pdf

Zhang, R., Song, G., Qu, Y., Alström, P., Ramos, R., Xing, X., Ericson, P.G.P., Fjeldså, J., Wang, H., Yang, X., Kristin, A., Shestopalov, A.M., Choe, J.C. & Lei, F. 2012. Comparative phylogeography of two widespread magpies: Importance of habitat preference and breeding behavior on genetic structure in China. Molecular Phylogenetics and Evolution 65: 562–572. DOI: 10.1016/j.ympev.2012.07.011 [Abstract] pdf

Fregin, S., Haase, M., Olsson, U. & Alström, P. 2012. Pitfalls in comparisons of genetic distances: A case study of the avian family Acrocephalidae. Molecular Phylogenetics and Evolution 62: 319–328.   DOI:10.1016/j.ympev.2011.10.003 [Abstract] pdf

Alström, P., Höhna, S., Gelang, M., Ericson, P.G.P. & Olsson, U.  2011. Non-monophyly and intricate morphological evolution within the avian family Cettiidae revealed by multilocus analysis of a taxonomically densely sampled dataset. BMC Evolutionary Biology 11: 352.   DOI:10.1186/1471-2148-11-352 [Open access] pdf

Ödeen, A, Håstad, O. & Alström, P. 2011. Evolution of ultraviolet vision in the largest avian radiation - the passerines. BMC Evolutionary Biology 11: 313.   DOI:10.1186/1471-2148-11-313 [Open access] pdf

Alström, P., Saitoh, T., Williams, D., Nishiumi, I., Shigeta, Y., Ueda, K., Irestedt, M., Björklund, M. & Olsson, U. 2011. The Arctic Warbler Phylloscopus borealis – three anciently separated cryptic species revealed. Ibis 153: 395–410.   DOI: 10.1111/j.1474-919X.2011.01116.x [Abstract] [Additional file] pdf

Alström, P., Fregin, S., Norman, J.A., Ericson, P.G.P., Christidis, L. & Olsson, U. 2011. Multilocus analysis of a taxonomically densely sampled dataset reveal extensive non-monophyly in the avian family Locustellidae. Molecular Phylogenetics and Evolution 58: 513–526.   DOI:10.1016/j.ympev.2010.12.012 [Abstract] [Figure 1] [Figure 2] [Figure 3] pdf

Irestedt, M., Gelang, M., Olsson, U., Ericson, P.G.P., Sangster, G. & Alström, P. 2011. Neumann’s Warbler Hemitesia neumanni (Sylvioidea): the sole African member of a Palaeotropic Miocene radiation. Ibis 153: 78–86.   DOI: 10.1111/j.1474-919X.2010.01084.x pdf

Alström, P., Fjeldså, J., Fregin, S. & Olsson, U. 2011. Gross morphology betrays phylogeny – the Scrub Warbler Scotocerca inquieta is not a cisticolid. Ibis 153: 87–97.   DOI: 10.1111/j.1474-919X.2010.01093.x pdf

Sangster, G., Alström, P., Forsmark, E. & Olsson, U. 2010. Multilocus phylogenetic analysis of Old World chats and flycatchers reveals extensive paraphyly at family, subfamily and genus level (Aves: Muscicapidae). Molecular Phylogenetics and Evolution 57: 380–392.   DOI:10.1016/j.ympev.2010.07.008 [Abstract] [Fig. 1] [Table 1] [Table 2] pdf

Leader, P.J., Carey, G.J., Olsson, U., Baral, H.S. & Alström, P. 2010. The taxonomic status of Rufous-rumped Grassbird Graminicola bengalensis, with comments on its distribution and status. Forktail 26: 121–126. pdf

Saitoh, T., Alström, P., Nishiumi, I., Shigeta, Y., Williams, D., Olsson, U. & Ueda, K. 2010. Old divergences in a boreal bird supports long-term survival through the Ice Ages. BMC Evolutionary Biology.   DOI:10.1186/1471-2148-10-35 [Open access] pdf

Alström, P., Davidson, P., Duckworth, J.W., Eames, J.C., Le, T.T., Nguyen, C., Olsson, U., Robson, C. & Timmins, R.J. 2010. Description of a new species of Phylloscopus warbler from Vietnam and Laos. Ibis 152: 145–168.   DOI: 10.1111/j.1474-919X.2009.00990.x [Abstract] [Photos] [Sound recordings] pdf

Olsson, U., Alström, P., Svensson, L., Aliabadian, M. & Sundberg, P. 2009. The Lanius excubitor (Aves, Passeriformes) conundrum – Taxonomic dilemma when molecular and non-molecular data tell different stories. Molecular Phylogenetics and Evolution 55: 347–357.   DOI:10.1016/j.ympev.2009.11.010. [Abstract] [Fig. 2] pdf

Ödeen, A., Håstad, O. & Alström, P. 2009. Evolution of ultraviolet vision in shorebirds (Charadriiformes). Biology Letters 6: 370–374.   DOI: 10.1098/rsbl.2009.0877. [Open accesspdf

Fregin, S., Haase, M., Olsson, U. & Alström, P. 2009. Multi-locus phylogeny of the family Acrocephalidae (Aves: Passeriformes) – The traditional taxonomy overthrown. Molecular Phylogenetics and Evolution 52: 866–878.   DOI:10.1016/j.ympev.2009.04.006 [Abstract] [Figure 3pdf

Gelang, M., Cibois, A., Pasquet, E., Olsson, U., Alström P. & Ericson, P.G.P. 2009. Phylogeny of babblers (Aves, Passeriformes): major lineages, family limits and classification. Zoologica Scripta 35: 225–236. pdf

Alström, P, Rasmussen, P.C., Olsson, U., & Sundberg, P. 2008. Species delimitation based on multiple criteria: the Spotted Bush Warbler Bradypterus thoracicus complex (Aves, Megaluridae). Zoological Journal of the Linnean Society 154: 291–307. pdf

Olsson, U., Alström, P., Gelang, M., Ericson, P.G.P. & Sundberg, P. 2008. What is proper vouchering in phylogenetic studies of birds? – a reply to Peterson et al. (2007). Molecular Phylogenetics and Evolution 48: 383–385.   DOI:10.1016/j.ympev.2008.03.031 pdf

Alström, P., Olsson, U., Lei, F., Wang, H-t., Gao, W. & Sundberg, P. 2008. Phylogeny and classification of the Old World Emberizini (Aves, Passeriformes). Molecular Phylogenetics and Evolution 47: 960–973.   DOI:10.1016/j.ympev.2007.12.007 [Abstract] [Fig. 3pdf

Nylander, J.A.A., Olsson, U., Alström, P. & Sanmartín, I. 2008. Accounting for phylogenetic uncertainty in biogeography: a Bayesian approach to dispersal-vicariance analysis of the thrushes (Aves: Turdus). Systematic Biology 57: 257–268. pdf [Suppl. fig.] [Suppl. fig. caption] [Suppl. table]

Johansson, U., Alström, P., Olsson, U., Ericson, P.G.P., Sundberg, P. & Price, T.D. 2007. Build-up of the Himalayan avifauna through immigration: a biogeographical analysis of the Phylloscopus and Seicercus warblers. Evolution 61: 324–333.   DOI: 10.1111/j.1558-5646.2007.00024.x [Abstract] [Fig. 2] [Fig. 3] pdf

Alström, P, Olsson, U., Rasmussen, P.C., Yao, C-t, Ericson, P.G.P. & Sundberg, P. 2007. Morphological, vocal and genetic divergence in the Cettia acanthizoides complex (Aves: Cettiidae). Zoological Journal of the Linnean Society 149: 437–452. pdf

Olsson, U., Alström, P., Gelang, M., Ericson, P.G.P. & Sundberg, P. 2006. Phylogeography of Indonesian and Sino-Himalayan region bush warblers (Cettia, Aves). Molecular Phylogenetics and Evolution 41: 556–565.   DOI:10.1016/j.ympev.2006.05.009 [Abstract] [Fig. 2] pdf

Alström, P. 2006. Species concepts and their application: insights from the genera Seicercus and Phylloscopus. Proceedings of the 23rd International Ornithological Congress, Acta Zoologica Sinica 52: 429–434. pdf

Alström, P, Ericson, P.G.P., Olsson, U., & Sundberg, P. 2006. Phylogeny and classification of the avian superfamily Sylvioidea. Molecular Phylogenetics and Evolution 38: 381–397.   DOI:10.1016/j.ympev.2005.05.015 [Abstract] [Fig. 3] pdf

Olsson, U., Alström, P., Ericson, P.G.P. & Sundberg, P. 2005. Non-monophyletic taxa and cryptic species – evidence from a molecular phylogeny of leaf-warblers (Phylloscopus, Aves). Molecular Phylogenetics and Evolution 36: 261–276.   DOI:10.1016/j.ympev.2005.01.012 [Abstract] [Fig. 1] [Fig. 5] [Fig. 6] [Fig. 7] [Table 1] pdf

Duckworth , J.W. & Alström, P. 2004. Motacilla samveasnae is the correct scientific name for the Mekong Wagtail. Bulletin of the British Ornithologists’ Club 124: 290–292. pdf

Olsson, U., Alström, P. & Sundberg, P. 2004. Non-monophyly of the avian genus Seicercus (Aves: Sylviidae) revealed by mitochondrial DNA. Zoologica Scripta 33: 501–510. pdf

Alström, P. & Ranft, R. 2003. The use of sounds in avian systematics, and the importance of bird sound archives. Bulletin of the British Ornithologists’ Club Supplement 123A: 114–135. pdf

Duckworth, J.W., Alström, P., Davidson, P., Evans, T.D., Poole, C.M., Tan S. & Timmins, R.J. 2001. A new species of wagtail from the lower Mekong basin. Bulletin of the British Ornithologists’ Club 121: 152–182. pdf

Irwin, D., Alström, P., Olsson, U. & Benowitz-Fredericks, Z.M. 2001. Cryptic species in the genus Phylloscopus (Old World leaf warblers). Ibis 143: 233–247.   DOI: 10.1111/j.1474-919X.2001.tb04479.x [Abstract] [Fig. 4] pdf

Alström, P. & Olsson, U. 2000. Golden-spectacled Warbler systematics. Ibis 142: 495–500.   DOI: 10.1111/j.1474-919X.2000.tb04449.x pdf

Alström, P. & Olsson, U. 1999. The Golden-spectacled Warbler: a complex of sibling species, including a previously undescribed species. Ibis 141: 545–568.   DOI: 10.1111/j.1474-919X.1999.tb07363.x [Abstract] [Fig. 4] [Fig. 5] [Fig. 6] [Fig. 7] [Fig. 12] [Table 1] [Table 2] [Table 3] [Table 4pdf

Alström, P. 1998. Taxonomy of the Mirafra assamica complex. Forktail 13: 97–107. pdf

Alström, P., Olsson, U. & Colston, P. 1997. Re-evaluation of the taxonomic status of Phylloscopus proregulus kansuensis Meise. Bulletin of the British Ornithologists’ Club 117: 177–193. pdf

Alström, P. & Olsson, U. 1995. A new species of Phylloscopus warbler from Sichuan Province, China. Ibis 137: 459–468.   DOI: 10.1111/j.1474-919X.1995.tb03254.x [Abstract] [Photo]

Alström, P., Olsson, U. & Colston, P.R. 1995. Revaluation of the taxonomic status of Phylloscopus ricketti goodsoni Hartert. Bulletin of the British Ornithologists’ Club 115: 53–57.

Alström, P. & Olsson, U. 1993. Blyth’s Leaf Warbler Phylloscopus reguloides found breeding in Thailand. Forktail 9: 150–152.

Alström, P., Ripley, S. D. & Rasmussen, P. C. 1993. Re-evaluation of the taxonomic status of Phylloscopus subaffinis arcanus. Bulletin of the British Ornithologists’ Club. 113: 207–209.

Alström, P. & Mild, K. 1993. Taxonomic status of Anthus berthelotii. Bulletin of the British Ornithologists’ Club 113: 88–92.

Olsson, U., Alström, P. & Colston, P.R. 1993. A new species of Phylloscopus warbler from Hainan Island, China. Ibis 135: 2–7.   DOI: 10.1111/j.1474-919X.1993.tb02803.x [Abstract] [Photo]

Alström, P., Olsson, U. & Colston, P. R. 1992. A new species of Phylloscopus warbler from central China. Ibis 134: 329–334.   DOI: 10.1111/j.1474-919X.1992.tb08011.x [Abstract] [Photo 1] [Photo 2]

Alström, P. & Olsson, U. 1992. Taxonomic status of Phylloscopus affinis and P. subaffinis. Bulletin of the British Ornithologists’ Club 112: 111–126. pdf

Alström, P., Olsson, U. & Round, P.D. 1991. The taxonomic status of Acrocephalus agricola tangorum. Forktail 6: 3–13.

Alström, P., Olsson, U. & Colston, P. R. 1990. Description of a possible new species of leaf warbler of the genus Phylloscopus from China. Bulletin of the British Ornithologists’ Club 110: 43–47.

Alström, P. & Olsson, U. 1990. Taxonomy of the Phylloscopus proregulus complex. Bulletin of the British Ornithologists’ Club 110: 38–43.

Alström, P. & Schekkerman, H. 1988. Desert Wheatear kleptoparasitizing ants. Sandgrouse 10: 111.

Alström, P. & Olsson, U. 1988. Gynandromorphism in the Pink-browed Rosefinch. Bulletin of the British Ornithologists’ Club 108: 12.

Books

Alström, P. & Mild, K. 2003. Pipits and Wagtails of Europe, Asia and North America: identification and systematics. London: Christopher Helm/A&C Black; Princeton: Princeton University Press. 496 pp. [Selected chapters from Introduction] [Richard's Pipit text] [Richard's Pipit plate]

Alström, P. & Colston, P. 1991. Field Guide to the Rare Birds of Britain and Europe. London: HarperCollins. 448 pp.

Book chapters

Alström, P. & Källersjö, M. 2007. Vem är släkt med vem? En inblick i systematikerns laboratorium. Pp. 154–175 in Från Linné till DNA. Stockholm: Naturhistoriska riksmuseet. [In Swedish with English summary] pdf

Bairlein, F., Alström, P., Aymí, R., Clement, P., Dyrcz, A., Gargallo, G., Hawkins, F., Madge, S., Pearson, D. & Svensson, L. 2006. Family Sylviidae (Warblers). Pp. 492–709 in del Hoyo, J., Elliott, A. & Christie, D.A. (Eds.). Handbook of the Birds of the World. Vol. 11. Barcelona: Lynx Edicions.

de Juana, E., Suárez, F., Ryan, P., Alström, P. & Donald, P. 2004. Family Alaudidae (Larks). Pp. 496–601 in del Hoyo, J., Elliott, A. & Christie, D.A. (Eds.). Handbook of the Birds of the World. Vol. 9. Barcelona: Lynx Edicions.

Abstracts in conference books

Gelang, M., Cibois, A., Pasquet, E., Mohammad, I., Olsson, U., Alström, P.& Ericson, P.G.P. 2010. Neogene biogeography of babblers. 25th International Ornithological Congress, Campos do Jordao, Brazil.

Gelang, M., Cibois, A., Pasquet, E., Olsson, U., Alström, P.& Ericson, P.G.P. 2009.Miocene and Pliocene radiations and biogeography of Leiothrichinae babblers (Aves, Timaliidae). Biennial meeting of International Biogeography Society, Merida,

 Alström, P., Olsson, U., Gelang, M., Rasmussen, P.C., Ericson, P.G.P. & Sundberg, P. 2006. Divergence in morphology, vocalisations and behaviour in a clade of Old World warblers. 24th International Ornithological Congress, Hamburg, Germany.

Saitoh, T., Nishiumi, I., Alström, P., Olsson, U. & Ueda, K. 2006. Deep phylogenetic divergences among Far Eastern populations of the widespread Arctic Warbler Phylloscopus borealis. 24th International Ornithological Congress, Hamburg, Germany.

Alström, P., Olsson, U., Rasmussen, P.C. & Sundberg, P. 2005. Song divergence and sympatry among taxa of the Spotted Bush-warbler Bradypterus thoracicus complex. American Ornithologists’ Union Meetings, UC Santa Barbara.

Popular science papers

Alström, P. 2010. Mångfald – ett myller av arter. Biodiverse 15: 3. [In Swedish] pdf

Alström, P. 2009. Nya fågelarter. Museosaurien No. 1/2009. [In Swedish]

Alström, P. & Mild, K. 2005. Biologiska, fylogenetiska och monofyletiska arter – skillnader och likheter. Fåglar i Uppland 31: 12–21. [In Swedish] pdf

Alström, P. & Mild, K. 2004. ‘Biological’, ‘phylogenetic’ and ‘monophylectic’ species – same or different? Alula 3/2004: 96–103. pdf

Alström, P. 2004. Fältbestämning av öronskarv. Roadrunner 1/2004: 42–44. [In Swedish]

Alström, P. & Mild, K. 2003. Flava flow. Birdwatch 2003, pp. 30–32.

Alström, P. 2001. Identification of Pechora Pipit. Roadrunner 9: 55–65.

Alström, P. 1997. What is a species? Species concepts in ornithology. Vår Fågelvärld 8/1997, pp. 6–15. [In Swedish]

Alström, P. 1997. Field identification of Asian Gyps vultures. Bull. Oriental Bird Club No. 25, pp. 32–49. pdf

Alström, P. & Mild, K. 1997. Die Bestimmung des Steppenpiepers Anthus godlewskii. Limicola 11: 97–117. [In German with English summary]

Alström, P. & Mild, K. 1997. Die Bestimmung des Wasserpieper-Komplexes: Berg- Anthus spinoletta, Strand- A. petrosus, Pazifikpieper A. rubescens und Rosenpieper A. roseatus. Limicola 11: 1–24. [In German with English summary]

Alström, P. & Mild, K. 1996. The identification of Rock, Water and Buff-bellied Pipits. Alula 2: 161–175. 

Alström, P. & MacKay, A. 1995. Identification of Siberian, Black-throated, Radde’s, Arabian and Brown Accentors. Birding World 8: 108–112.

Alström, P., Olsson, U. & Round, P.D. 1994. Bestimmung der kleinen fernöstlichen Rohrsänger Acrocephalus. Limicola 8: 121–131. [In German with English summary]

Alström, P. & Olsson, U. 1994. Identification of Tickell’s Warbler Phylloscopus affinis and Buff-throated Warbler P. subaffinis. Dutch Birding 16: 89–94.

Alström, P. & Olsson, U. 1994. Identification of Pallas’s Reed Bunting Emberiza pallasi. Birding World 6: 15–20.

Alström, P. 1993. Common Gull Larus canus recorded in India. Journal of the Bombay Natural History Society 90: 509–510.

Alström, P. 1993. Understanding moult. Birding World 6: 198-205.

Alström, P. 1993. Identification of Bonaparte's Gull Larus philadelphia. Vår Fågelvärld 1/93, pp. 38–40. [In Swedish]

Alström, P. 1992. A Radde’s Accentor Prunella ocularis from Oman reidentified as Black-throated Accentor P. atrogularis. Sandgrouse 13: 106–108.

Alström, P. 1992. Identification of Long-billed and Short-billed Dowitchers. Fåglar i Sörmland 25: 19–25. [In Swedish]

Alström, P. 1991. Identification of Double-crested Cormorant. Birding World 4: 9–16.

Alström, P. 1991. Identification of Lesser Sand Plover. Vår Fågelvärld 50: 33–37. [In Swedish]

Alström, P. 1991. Identification of Catharus thrushes. Vår Fågelvärld 50: 58–60. [In Swedish]

Alström, P., Mild, K. & Zetterstrom, B. 1991. Identification of Lesser Short-toed Lark. Birding World 4: 422–427.

Alström, P. & Mild, K. 1991. Identification of Olive-backed Pipit. Vår Fågelvärld 50: 65–69. [In Swedish]

Alström, P. & Schekkerman, H. 1990. Desert Wheatear kleptoparasitizing ants. Sandgrouse 10: 111.

Shirihai, H. & Alström, P. 1990. Identification of Hume’s Short-toed Lark. British Birds 83: 262–272.

Alström, P. 1990. Calls of American and Pacific Golden Plovers. British Birds 83: 70–72.

Alström, P. & Mild, K. 1989. Identification of Pechora Pipit. Birding World 2: 276–282.

Alström, P. & Mild, K. 1989. Red-throated Pipits with rufous throat in winter. Dutch Birding 11: 78–81.

Alström, P. & Mild, K. 1989. Identification of marsh terns in juvenile and winter plumages. British Birds 82: 296–319.

Alström, P. & Olsson, U. 1989. The pattern on the sides of the lower mandible of Whooper Swan and Bewick's Swan. Vår Fågelvärld 48: 24. [In Swedish with English summary]

Alström, P. & Olsson, U. 1989. The identification of juvenile Red-necked and Long-toed Stints. British Birds 82: 360–372.

Alström, P. & Hirschfeld, E. 1989. Identification of three northeast Asian flycatchers in the genus Muscicapa. Vår Fågelvärld 48: 127–138. [In Swedish with English summary]

Alström, P. 1989. Identification of Lanceolated Warbler Locustella lanceolata. Vår Fågelvärld 48: 335–346. [In Swedish with English summary]

Alström, P. & Olsson, U. 1988. Taxonomy of Yellow-browed Warblers. British Birds 81: 656–657.

Alström, P. 1988. Identification of Blyth's Pipit. Birding World 1: 268–272.

Alström, P. & Mild, K. 1988. Calls of Blyth's Pipit. British Birds 81: 655.

Alström, P. & Mild, K. 1987. Some notes on the taxonomy of the Water Pipit complex. Proc. 4th Intern. Identification Meeting, p. 47–48. Eilat.

Alström, P. & Olsson, U. 1987. Field identification of Arctic and Greenish Warblers. Proc. 4th International Identification Meeting pp. 54–59. Eilat

Alström, P. & Olsson, U. 1987. Some rare species observed in Japan from September to November 1984. Strix 6: 105–108. [In Japanese with English summary]

Alström, P. 1986. Rare birds in Sweden in 1984 – report from the Swedish Rarities Committee. Vår Fågelvärld 45: 61–71. [In Swedish with English summary]

Alström, P. 1985. Identification of Cormorant Phalacrocorax carbo and Shag Ph. aristotelis. Vår Fågelvärld 44: 325–350. [In Swedish with English summary]

Alström, P. 1985. Identification of Baird's Sandpiper Calidris bairdii and White-rumped Sandpiper C. fuscicollis in juvenile plumage. Vår Fågelvärld 44: 479–484. [In Swedish with English summary]

Alström, P. & Olsson, U. 1985. First record of Iceland Gull Larus glaucoides from Japan. Strix 4: 70–72. [In Japanese with English summary]

Alström, P. 1983. First record of Alpine Swift Apus melba for Sweden. Vår Fågelvärld 42: 89–90. [In Swedish with English summary]

Alström, P. and Olsson, U. 1982. Bill colour of Crested Coot. British Birds 75: 287.

Alström, P. et al. 1981. Bird report from the Swedish west coast 1978-1979. Fåglar på Västkusten, supplement 2. [In Swedish]

Götmark, F., Wallin, K., Jacobsson, S. & Alström, P. 1979. Visible bird migration at Mönster in northern Halland, south-western Sweden, in the autumns of 1976 and 1977. Vår Fågelvärld 38: 201–220. [In Swedish with English summary]

Alström, P. et al. 1979. Bird report from the Swedish west coast 1976-1977. Fåglar på Västkusten, supplement 1. [In Swedish]

Alström, P. 1977. An influx of Parrot Crossbill. Fåglar på Västkusten 11: 39–42. [In Swedish]

Lectures etc.

 Lecture on biodiversity at Flora och faunavårdskonferensen, Uppsala, 2010 (in Swedish)

 Interview Sveriges Radio, Vetandets Värld, 24 April, 2012 (in Swedish)

 Interview Swedish Radio, Studio Uppland, 12 March, 2012 (in Swedish)

 Interview, Roadrunner 1-2011 (in Swedish) 

 Lecture on identification of Phylloscopus and Seicercus warblers, Beijing Birdwatching Society, November 2012. (English pdf) (Chinese pdf)

 Intervju i boken Skatter i vått och torrt (0,65 MB)

 
Sidan uppdaterad: 2014-05-22. Sidansvarig: per.alstrom@slu.se