Root physiologi, mycorrhizal symbiosis, plant nutrition.
- Plant physiology, plant nutrition, mycorrhizas, soil chemistry
- Project course in cooperation with the horticultural industry
- Course leader for Plant Physiology for Horticultural Management Programmes, Project course in Horticulture, Basics of Technological and Biology for Horticultural Production.
Master of Science in Plant Physiology 1988
PhD in Horticulture 1997