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Simon Kärvemo

The structure of the habitat and surrounding landscapes are crucial for the biodiversity and interaction among species. My research have mainly focused on landscape ecology, population dynamics and risk of infestations and infections of parasites. Current research questions involves forest structures and conservation areas and how it affect bark beetle infestations and biodiversity.


My research focus on factors affecting population dynamics of the spruce bark beetle Ips typographus across spatial scales, but I am also engaged in diverse projects including forest (and wetland) restorations, with a particular emphasis on understanding their implications for forest pests and biodiversity conservation but also the prevalence of amphibian infections. Presently, my main research areas involve the dynamics of bark beetle outbreaks, the efficacy of conservation areas, and how infestations affect forest biodiversity.



Mainly courses in entomology and herpetology (more information on the Swedish version)


- Bark beetles and forest conservation

- Woodland key habitats and effects on biodiversity 

- Distribution of the chytrid fungus in nordic amphibians

- Habitat choice and movements in reptiles and amphibians

- Forest restorations and effects on bark beetle diversity

- Outbreak dynamics of the spruce bark beetle Ips typographus 



Previous master/candidate students:

Maria Johansson, Sveriges lantbruksuniversitet (huvudhandledare)

Martin Westerling, Sveriges lantbruksuniversitet (huvudhandledare)

Evelina Andersson, Sveriges lantbruksuniversitet (huvudhandledare)

Patric Falck, Sveriges lantbruksuniversitet (huvudhandledare)

Ludvig Palmheden, Sveriges lantbruksuniversitet (huvudhandledare)

Filippa Hedenberg, Sveriges lantbruksuniversitet (biträdande handledare)

Malin Karlsson, Sveriges lantbruksuniversitet (biträdande handledare)

Gustav Wikström, Uppsala universitet (huvudhandledare)

David Jonsved, Uppsala universitet (huvudhandledare)

Anais Voisin, Stockholms universitet (biträdande handledare)

Gustaf Pettersson, Sveriges lantbruksuniversitet (huvudhandledare)

Are you looking for a masterproject? Bark beetles and occationally herpetological projects. Are you interested in GIS-analyses, spatial ecology, advanced R-coding or how landscaped are affecting pathogenes, pest insects or biodiveristy? Send an email to

Selected publications

For a complete list of publications, see my Google Scholar profile

Persson, H. J., Kärvemo, S., Lindberg, E., & Huo, L. (2024). Large-scale wall-to-wall mapping of bark beetle damage and forest practices using the distance red swir index and operational harvester data. Ecological Indicators, 162, 112036.

Elmberg, J., Palmheden, L., Edelstam, C., Hagman, M., & Kärvemo, S. (2024). Climate change-induced shifts in survival and size of the world's northernmost oviparous snake: A 68-year study. Plos one, 19(3), e0300363.

Kärvemo, S., Huo, L., Öhrn, P., Lindberg, E., & Persson, H. 2023. Different triggers, different stories: bark-beetle infestation patterns after storm and drought-induced outbreaks. Forest Ecology and Management. 

Hekkala, A. M., Jönsson, M., Kärvemo, S., Strengbom, J., & Sjögren, J. 2023. Habitat heterogeneity is a good predictor of boreal forest biodiversity. Ecological Indicators, 148: 110069.

Kärvemo, S., Schroeder, M. & Ranius, T. 2023. Beetle diversity in dead wood is lower in non-native tree species. Journal of Applied Ecology. 60:170-108

Multi-taxon conservation in northern forest hot-spots: the role of forest characteristics and spatial scales (2021). S. Kärvemo, M. Jönsson, AM. Hekkala, J. Sjögren & J, Strengbom. Landscape Ecology, 1-14.

Hekkala, A-M., Kärvemo, S., Versluijs, M., Weslien, J-O, Löfroth, T., Björkman, C. & Hjältén, J. (2020). Balancing conflicting goals: ecological restoration for biodiversity conservation and the response of bark beetles and their enemies.  Forestry 95: 115-126.

Kärvemo, S., Wikström, G., Widenfalk, L., Höglund, J & Laurila, A. (2020). Chytrid dynamics and infection trade-offs with movement distances in a red-listed amphibian. Journal of Zoology. 311: 164-174

Kärvemo, S., Laurila, A. & Höglund, J. (2019). Urban environment and reservoir host species are associated with  Batrachochytrium dendrobatidis infection prevalence in the common toad. Diseases of aquatic organisms. 34: 33-42

Kärvemo, S., Meurling, S., Berger, D., Höglund, J. & Laurila, A. (2018). Effects of host species and environmental factors on the prevalence of Batrachochytrium dendrobatidis in northern Europe. PLOS One 13(10): e0199852. https://doi.

Hallinger, M., Kärvemo, S., & Ranius, T. (2017). Does it pay to concentrate conservation efforts for dead-wood dependent insects close to existing reserves: a test on conservation planning in a Swedish region. Insect Conservation and Diversity. DOI: 10.1111/icad.12279

Kärvemo, S., Björkman, C., Johansson, T., Weslien, J-O, Hjältén, J. 2017. Forest restoration as a double-edged sword: the conflict between biodiversity conservation and pest control. Journal of Applied Ecology. 54: 1658-1668.

Kärvemo, S., Johansson, V., Schroeder, M. & Ranius, T. 2016. Colonisation-extinction dynamics of a tree-killing bark beetle. Ecosphere. 7:e01257.

Kärvemo, S., Rogell, B., & Schroeder, M. 2014. Dynamics of spruce bark beetle infestation spots: importance of local population size and landscape characteristics after a storm disturbance. Forest Ecology and Management. 334: 232-240.

Kärvemo, S., Van Boeckel, T.P., Gilbert, M., Grégoire, J-C., & Schroeder, M. 2014. Large-scale risk mapping of an eruptive bark beetle - Importance of forest susceptibility and beetle pressure. Forest Ecology and Management 318: 158-166.

Kärvemo, S., & Schroeder, L. M. 2010. A comparison of outbreak dynamics of the spruce bark beetle in Sweden and the mountain pine beetle in Canada (Curculionidae: Scolytinae). Entomologisk tidsskrift 131, nr 3, 215-242.